COLUMBUS, Ohio (AP) — Trustees of the Columbus Zoo and Aquarium said Friday that financial abuses by four former executives cost the institution more than $630,000, a new detail in the scandal that has marred the reputation of one of the nation’s largest zoos.

A forensic analysis ordered by the zoo’s board of trustees showed money collected through a zoo levy that pays for animal care not was involved, however.

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The review was ordered after a Columbus Dispatch investigation showed apparent improprieties by the zoo’s then-chief executive and chief financial officers, including misuse of zoo properties and sports tickets.

The review ordered by trustees found a former marketing vice president and purchasing director also were involved.

The full review, conducted by Plante Moran and overseen by Porter Wright Morris & Arthur, was delivered to key state, county and city officials and to the Ohio Ethics Commission on Friday. It will be released to the public Saturday.

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In highlights released Friday, the zoo said former president and CEO Tom Stalf was responsible for $423,000 of the losses, while former CFO Greg Bell was responsible for $139,000. Peter Fingerhut, then-vice president of marketing, was responsible for $57,000, while former director of purchasing Tracy Murname was responsible for $13,000.

“We trusted these individuals with the privilege to lead this singularly outstanding organization, and they shattered that trust,” board chairman Keith Shumate said in a statement.

He said the board and new leadership team will take immediate action to restore public trust. Trustees are reviewing recommendations made as part of the review.

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